Palaeontologia africana

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ISSN (print): 0078-8554 ISSN (electronic): 2410-4418 For queries regarding content of Palaeontologia africana collections please contact Jonah Choiniere by email : jonah.choiniere@wits.ac.za or Tel : 011 717 6684

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    Cranial morphology and phylogenetic analysis of Cynosaurus suppostus (Therapsida, Cynodontia) from the upper Permian of the Karoo Basin, South Africa
    (Evolutionary Studies Institute, 2018-03) van den Brandt, Marc; Abdala, Fernando
    Non-mammaliaform cynodonts are an important fossil lineage which include the ancestors of mammals and which illustrate the gradual evolution of mammalian characteristics. The earliest cynodonts (‘basal cynodonts’) are known from the late Permian. Cynosaurus suppostus is the second most abundant basal cynodont from the late Permian of the Karoo Basin of SouthAfrica, but is poorly studied, with the most recent description of this taxon being 50 years old. Since then, several important new specimens of this species have been collected, meriting a thorough description of its cranial anatomy and exploration of its interspecific variation. Here we present a comprehensive description of the cranial morphology of Cynosaurus suppostus, producing an updated diagnosis for the species and comparisons amongst basal cynodonts. Cynosaurus is identified by three autapomorphies amongst basal cynodonts: a subvertical mentum on the anterior lower jaw; a robust mandible with a relatively high horizontal ramus; and the broadest snout across the canine region, representing up to 31.74% of basal skull length. One of the new specimens described here preserves orbital scleral ossicles, structures rarely preserved in non-mammaliaform cynodonts. Cynosaurus is now only the third cynodont in which scleral ossicles have been reported.Anupdated phylogenetic analysis of basal cynodont interrelationships recovered Cynosaurus suppostus as a member of the Galesauridae in only two of 16 most parsimonious trees, providing poor support for its inclusion in that family. The majority of known Cynosaurus specimens were collected in a geographically restricted area approximately 150 kilometres in diameter. Most specimens have been recovered from the latest Permian Daptocephalus Assemblage Zone, with only two specimens known from the older Cistecephalus Assemblage Zone.
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    A review of the reptile and amphibian assemblages from the Stormberg of southern Africa, with special emphasis on the footprints and the age of the Stormberg
    (Bernard Price Institute for Palaeontological Research, 1984) Olsen, Paul E.; Galton, Peter M.
    The Molteno, Elliot, and Clarens formations comprise the continental Stormberg Group of the Karoo Basin of South Africa and Lesotho. The Molteno Formation contains a well preserved macro- and microfloral assemblage but apparently no vertebrates; the Elliot and Clarens formations contain abundant vertebrates but virtually no floral remains. The vertebrate taxa represented by skeletal remains are listed and divided into two assemblages - the lower Stormberg (lower Elliot) and upper Stormberg (upper Elliot and Clarens) assemblages. The abundant, diagnosable footprint taxa are revised and their names reduced to eight genera. These ichnotaxa also fall into two biostratigraphic zones that parallel the skeletal assemblages. Comparison of the faunal assemblages with those of the European type section strongly suggests that the lower Stormberg assemblage is Late Triassic (Carnian- Norian) in age while the upper Stormberg assemblage is Early Jurassic (Hettangian-Pliens- bachian) in age. Comparisons with other continental assemblages from other areas suggest that the upper Stormberg (upper Elliot and Clarens formations) assemblage broadly correlates with the upper Newark Supergroup of eastern North America, the Glen Canyon of the southwestern United States, and the lower Lufeng Series of China- all thought to be of Early Jurassic age on the basis of floral and/or radiometric evidence. Based on these correlations, previously published paleobiogeographic maps are revised; these show a shift from Late Triassic floral and faunal provinciality to Early Jurassic homogeneity. This shift was synchronous with a widening of the equatorial arid zone.
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    A structural re-interpretation and revision of the type material of the glossopterid ovuliferous fruitification Scutum from South Africa
    (Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, 2011-12) Prevec, Rose
    The Early Permian glossopterid fructification Scutum, described by Edna Plumstead in the 1950s from the Vereeniging locality in the Karoo Basin of South Africa, was one of the first glossopterid seed-bearing organs to be found in organic attachment to Glossopteris leaves. Examination of the type material necessitated a revision of this plant fossil genus and a re-evaluation of described South African species. Key characteristics of the genus are the broad and prominent wing, and a low receptacle length to width ratio (<2:1). Specimens of South African Scutum are currently attributed to three species, from two localities, but display intergrading morphological features that can be reasonably accommodated within a single species, S. leslii. Three-dimensional interpretation and reconstruction of impression fossils of Scutum fructifications preserved in attachment to Glossopteris leaves confirms that the seed-bearing surface of the receptacle faces the adaxial surface of the subtending leaf. The nature of the seed scars on the receptacle and their relationship to the peripheral wing of the fructifications is clarified.
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    Cranial morphology and taxonomy of South African Tapinocephalidae (Therapsida: Dinocephalia): the case of Avenantia and Riebeeckosaurus
    (Evolutionary Studies Institute, University of the Witwatersrand, 2013-12-18) Guven, Saniye; Rubidge, Bruce S.; Abdala, Fernando
    Tapinocephalid dinocephalians comprise a large component of Middle Permian herbivorous tetrapod biodiversity and are significantly abundant in the Karoo rocks of SouthAfrica. In order to understand the effect of the alleged Middle Permian extinction in the terrestrial record of amniotes a clear understanding of the taxonomic diversity of this group is essential. The tapinocephalids Avenantia kruisvleiensis and Riebeeckosaurus longirostris have in the past been distinguished by a set of characters, including differences in the snout length which was previously interpreted as a key diagnostic character. Further preparation of the material attributed to these species shows that the purported distinguishing characters, such as snout length, low intertemporal crest, and surface thickening of the parietal represent postmortem deformation (including distortion and weathering) or can be interpreted as individual or ontogenetic variation. Accordingly A. kruisvleiensis is synonimized with R. longirostris.