Palaeontologia africana

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Taxonomic and taphonomic interpretations of newly excavated in situ GD 2 faunal remains at Gondolin
(The Evolutionary Studies Institute, 2024-01) Engelbrecht, Micke; Val, Aurore; Kibii, Job M.; Steininger, Christine
Gondolin is one of the fossil-bearing karstic localities in the Cradle of Humankind, South Africa. Periodic excavations of calcified and decalcified sediments at several loci (GD 1,GD2 andGDA) over the last few decades have yielded a sizeable sample of Plio-Pleistocene fauna, including two hominin teeth from ex situ deposits. In 2015, renewed excavations were conducted at the GD 2 locality, which consists of decalcified in situ deposits, in order to shed more light on the site’s complex formation processes as well as to try finding new hominin material from a stratigraphically secured context. While these excavations did not yield any hominin material, abundant macrovertebrate remains were recovered. This paper presents the taxonomic composition and taphonomic characteristics of this new faunal assemblage. The occurrence of Equus sp. as well as the extinct species Metridiochoerus andrewsi and Hystrix makapanensis places the assemblage in a depositional age bracket of 2.33–1.78 Ma, which is consistent with ages already proposed forGD2. No primate material was recovered. The assemblage displays a taxonomic and bodyweight bias towards small-bodied (size classes I and II) bovids. The taphonomic characteristics of the bovid remains suggest the selective action of a leopard-like carnivore and while there is no direct evidence that the locality was used as a hyaena den, secondary scavenging by hyaenids cannot be excluded. Porcupines played an ancillary role in the bone accumulation. We performed intra- and inter-site taxonomic and taphonomic comparisons between this faunal sample and fossil assemblages from previously excavated localities at Gondolin (GD 1, GD 2 and GD A), as well as from other Paranthropus robustus-bearing sites in the Cradle of Humankind. These comparisons indicate that this new sample closely resembles the faunal assemblage previously collected from in situ calcified sediments at the GD 2 locality, in terms of species composition, bovid size class distribution, carnivore to ungulate ratio, and general taphonomic characterization.
The pre-eminence of the Karoo Basin in the knowledge of the Permo-Jurassic cynodonts: a historical synthesis and taxonomical quantification
(Evolutionary Studies Institute, 2023) Abdala, Fernando; Hendrickx, Christophe; Jasinoski, Sandra C; Gaetano, Leandro C; Liu, Jun
The search for the ancestors of mammals is historically connected with the extensive Karoo Basin of South Africa. This is because the Karoo features some of the largest exposures of Permo-Jurassic terrestrial deposits in the world and fossil discoveries were made here early in the history of palaeontology. Among the mammal-like lineages that are well-represented in Karoo fossil assemblages are the cynodonts. Originally conceived as a group exclusively containing fossil taxa, Cynodontia was subsequently redefined to include living mammals, and its Permian and early Mesozoic members are now referred to as non-mammaliaform cynodonts. Here we present a historical account of the research programme on non-mammaliaform cynodonts in the Karoo Basin, which represent the most important record of this group in the world. It covers a time spanning from the first named species in 1859 until the present day, which we arbitrarily divided into three periods: the Early Period extending from 1859 until 1932, the Second Period from 1933 to 1982, and the Current Period from 1983 until now. In the context of the global record of named species, we present quantitative analyses documenting the total number of nominal non-mammaliaform cynodont species from the Karoo (including junior synonyms and homonyms) as well as numerical comparison with taxa currently considered valid. Lastly, we compare the record of non-mammaliaform cynodont species from South Africa with other places in the world, such as Argentina and Brazil, which also have a diverse record of this group.

Palaeontologia africana

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